Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae)

Individuals of Boophis kirki sp. nov. in life. A, D Male paratype KU 336967 (CRH 20) from Ranomafana in dorsolateral and ventral view. B, E Male paratype UADBA-CRH 15 (CRH 15) from Ranomafana in dorsolateral and ventral view. C Male paratype KU 336874 (CRH 17) from Ranomafana. F Male paratype KU 336875 (CRH 18) from Ranomafana. G Female (no type status) UADBA-CRH 430 (CRH 430) from Vohidrazana. H Male (no type status) KU 340727 (CRH 449) from Vohidrazana. I, J Male (no type status) KU 340730 (CRH 455) from Vohidrazana in dorsolateral and ventral view. Note the variation in iris color in both populations (Ranomafana and Vohidrazana), with some individual having red color in the outer iris area while others only have light orange color.