Maximum credibility chronogram (MCC) inferred in BEAST v. 1.10.4 with bars, showing the 95% HPDI of divergence time estimates. The vertical gray line marks the Messinian Salinity Crisis (MSC; 5.96–5.33 Mya). Red circles at the nodes indicate used calibration points, white circles indicate posterior probabilities above 0.95, and black circles denote groups of taxa forced to be monophyletic in the analysis.